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Spice and Wolf's story revolves around Lawrence Craft, a twenty-five-year-old merchant. His main goal in life is to gather enough money to start his own shop, and he already has been traveling for seven years while gaining experience in the trade. One night when stopped at the town of Pasroe, he finds a 250-year-old pagan wolf-deity girl named Horo sleeping in his cart. She appears to be a fifteen-year-old girl, except she possesses a wolf's tail and ears. She introduces herself as the town's harvest goddess, who has kept the area blessed with good wheat crops for many years. Despite having the responsibility to watch over the town, she wishes to go back to her homeland in the north called Yoitsu, since she believes that the locals have forsaken their past agreement and therefore she is no longer bound to their land. She makes a deal with Lawrence to take her with him in exchange for her wisdom and supernatural abilities, but as they travel, her true nature draws unwanted attention from the church.
The protagonist of the series is the monk-in-training Ikkou Satonaka, who transforms into a super-monk (being able to perform mass exorcisms) by his lust for the girls he lives with (Note: In the anime, he transforms from seeing a naked girl). He lives in the Saienji Temple as a Buddhist priest in training with six other nuns: Haruka Amanogawa, Sumi Ikuina, Hinata and Sakura Sugai, Chitose Nanbu and Yuuko Atouda, each of whom represents one of the bosatsu of the six lower realms of the traditional Buddhist cosmology.